Joseph M. DiTomaso, University of California - Davis, Bugwood.org

Joseph M. DiTomaso, University of California - Davis, Bugwood.org

Tony Pernas, USDI National Park Service, Bugwood.org

Joseph M. DiTomaso, University of California - Davis, Bugwood.org

Joseph M. DiTomaso, University of California - Davis, Bugwood.org

Utah State University , Bugwood.org